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Stephen Jay Gould's popular 1989 account of this work, Wonderful Life, brought the matter into the public eye and raised questions about what the explosion represented.

However, life in these areas is becoming increasingly difficult, particularly in recent years due to the increasing scarcity of water.

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Whittington and colleagues, who, in the 1970s, reanalysed many fossils from the Burgess Shale and concluded that several were as complex as, but different from, any living animals.

The most common organism, Marrella, was clearly an arthropod, but not a member of any known arthropod class.

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